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Permian Brachiopods of West Texas, III


G. Arthur Cooper and Richard E. Grant

isSUED DEG 99 185


JAN 15 arb



Cooper, G. Arthur, and Richard E, Grant. Permian Brachiopods of West Texas, III. Smithsonian Contributions to Paleobiology, number 19 (part 1: text; part 2: plates), pages 795-1921, plates 192-502, 1975.—The third of a six-part mono- graph on the Permian brachiopods of the Glass, Guadalupe and other mountain ranges of West Texas, this volume contains systematic descriptions of genera and species in the suborders Productidina and Chonetidina. The Productidina, which constitute about 45 percent of the brachiopod specimens in the collections from West Texas, are divided into the superfamilies Strophalosiacea, Aulostegacea, Richthofeniacea, and Productacea. The Chonetidina, less numerous, contain the single superfamily Chonetacea.

Permian Brachiopods of West Texas, I. Smithsonian Contributions to Paleobiology, number 14, 231 pages, 39 figures, 23 plates. Issued 29 December 1972.

Permian Brachiopods of West Texas, II. Smithsonian Contributions to Paleobiology, number 15, pages 233-793, figure 40, plates 24-191. Issued 16 April 1974.

OFFICIAL PUBLICATION DATE is handstamped in a limited number of initial copies and is re- corded in the Institution’s annual report, Smithsonian Year. SI Press NUMBER 4808. SERIES COVER DESIGN: The trilobite Phacops rana Green.

Library of Congress Cataloging in Publication Data

Cooper, Gustav Arthur, .1902-

Permian brachiopods of West Texas.

Smithsonian contributions to paleobiology, no. 14-15, 19) upt. of Docs: SI 1.30: 19

Includes bibliographies.

1, Brachiopods, Fossils. 2. Paleontology—Permian. 3. Paleontology—Texas. I. Grant, Richard E., joint author. II, Title. III. Series: Smithsonian Institution. Smithsonian contributions to paleobiology, no. 14 [etc.]

QE701.856 no. 14, etc. [QE796] 560’.8s [564’.8'097649] 72-4218

For sale by the Superintendent of Documents, U.S. Government Printing Office, Washington, D.C. 20402 Price domestic postpaid or GPO Bookstore

Order Strophomenida Suborder Productidina

Superfamily Strophalosiacea ..

Family Strophalosiidae


Subfamily Strophalosiinae ..

Genus Strophalosia Genus Heteralosia Subfamily Ctenalosiinae . Genus Ctenalosia Family Teguliferinidae Genus Teguliferina Genus Acritosia Superfamily Aulostegacea_ .. Family Cooperinidae Genus Cooperina Genus Atelestegastus . Family Aulostegidae Subfamily Echinosteginae Genus Limbella Genus Edriosteges Genus Echinosteges .. Genus Cactosteges . Genus Xenosteges Subfamily Chonosteginae Genus Chonosteges .. Subfamily Institellinae .. Genus Institella Genus Glyptosteges .. Genus Craspedona Family Spyridiophoridae .. Genus Spyridiophora Family Agelesiidae Genus Agelesia ... .. Family Scapharinidae Genus Scapharina Family Rhamnariidae Genus Spuriosa Genus Ramavectus Genus Rhamnaria Family Tschernyschewiidae Genus Tschernyschewia Family Scacchinellidae Genus Scacchinella Superfamily Richthofeniacea Family Hercosiidae Genus Hercosia Genus Hercosestria Family Cyclacanthariidae .. Genus Cyclacantharia Genus Taphrosestria Genus Sestropoma Genus Collumatus Family Richthofeniidae ..


Genus Richthofenia Subfamily Prorichthofeniinae Superfamily Productacea Family Overtoniidae . Subfamily Overtoniinae .... Genus Fimbrinia Genus Simplicarina Genus Rhytisia Family Marginiferidae .. Subfamily Marginiferinae Genus Kozlowskia Genus Hystriculina .... .. Subfamily Costispiniferinae Genus Bothrionia Genus Costispinifera Genus Elliottella Genus Echinauris

Genus Oncosarina .........

Subfamily Retariinae Genus Kutorginella Genus Thamnosia

Family Echinoconchidae

Subfamily Echinoconchinae ....

Genus Echinaria .. ............

Genus Calliprotonia Genus Bathymyonia Subfamily Waagenoconchinae Genus Waagenoconcha Family Buxtoniidae .......... Subfamily Buxtoniinae Genus Kochiproductus .. Genus Ametoria .... Subfamily Juresaniinae Genus Juresania .. fae Family Dictyoclostidae .. ... Subfamily Dictyoclostinae .. Genus Spinarella Genus Xestosia yo os. Genus Nudauris Genus Dasysaria Genus Rugatia ........ Genus Spinifrons ..... ... Genus Reticulatia Genus Antiquatonia Genus Peniculauris Subfamily Horridoniinae .. Genus Horridonia Family Paucispiniferidae Genus Anemonaria Genus Liosotella Genus Paucispinifera Genus Polymorpharia Family Linoproductidae Subfamily Linoproductinae Genus Genus Cancrinella Genus Liraria Genus Undellaria Genus Grandaurispina Genus Holotricharina Genus Yakovlevia

Linoproductus ....,



Genus Siphonosia sce - os Sas iva vues have Pe che hoe Subfamily Anidanthinae .................... Fs pate viD Leach ies 2 a ecied Genus Mera sa Ord 5 Pin da alen ite ae ae el obiee day ee dla: Sublamily “Siviatiferinae .. meas. yas here oh Me cs atk he det re Genus JStriatiferd® 2 a hhee heb eesti yt Baia habs Re ete s thle ghee: Genus Compressoproductus —............ eS Bete Sg here itd QUE ere ase Uncertain and Misidentified Productacea .. 200 1 w. eee cece eee ees Suborder “Ghionetidina: qx ries y ate ale seih ed agey epee Dein Or AES eh chatted ye seta Superfamily Chonetacea ..... .... ........5. Bal eoetiregp rote styl al RO REALL pita ae Family Rugosochonetidae .. ...........0 ..... reba Ceres peepee Eee tee le Subfamily Rugosochonetinae .. .. ......... a xt Honah ia sy Genis: Siuleatayin 8 i Aw. Seehvconts boys D1 yard acest LM a ALR a Genus Neochonetes ..._....... Pht bee Bags Peete OM Lm ERE an baie Genus Dyoros (Dyoros) oi visite gen See ean aided als Genus Dyoros (Lissosia) .. ... sk AMR ES ba eet Bie at sa tecny Genus Dyoros (Tetragonetes) ..... ...... $e a st, Genus Lissochonetes ...............4.. Pf As) Peet: & bey og! Genus. Dewrosine: oso. fd edn amu ee BA ee een BOE Gerig MesoloGus Vu 2. 05 ee yee nls ely aeiating 7 @ LEY ostinnse Pie Feud Mess yp. Ye GOnUs “Griadrog none tess. 363 a6 lve ied Bere 5 AEE Rho ee SES ape hier iit 008 Subfamily Lamellosiinae Pe EE ere BR I a op seca! f Hube Mie 2 obs A Bs fe Genus Lamellosia .. 1.0.2... eee kit eS outaeeg ale, Sestak Subfamily “Chohetinellitiag tcc a ig tics nes shy the als te a oo sarenb 8 a da SONS GLOMETIMEL A ye tA Sh ee ito cs on Mite erat glee B etpde sty ek Rak Genus -Chonetinstes ee bel babies ging - dent Bl phn a as Subfamily Undulellinae ............. OT ee ae ere Penh eee | a Genus Undulella .....0 0 ......0 0 ween eae Se cd ee, tt etch one Genus Micraphelia __.................. oft) See Free See a Bo Subfamily Plicochonetinae ........... Genus Ftugaria on. eked hea petite able sie leltchty atalfiase Le thee te whe Literature iGited> ey) gO dh eh cntute ear gee tomes so. e ety See pet


192. Heteralosia, Ctenalosia, and Liosotella .....0 1.2.00 1. oe cc cee AOS: SEERETALOSED™ 938 ha. Ns ripen GAG eae Peta oe ideo ub es Vo wk ate ed eat 194, epuriosa- anu tileteralasial. 25 19 oa gh st tS nce cette aw oys bt ce aat oe Sree mas 195. Heteralosia ..... Meee tees lan Al yan siete nd Pau hy eye eas el release st tah ap ene 196. Heteralosia ..... rena a wey eed OR een pe bois Tinsel bee ees 197. Ctenalosia .. .. SS To AD EEN, NASR Fae cat ff An uate riots rte tig relay, beats RUSS ClenalOsta;* chen ig leet || BAR A ies, bq Sve) on eg Cea Ay at et pret enti TOO. AETHOSES NEG Keine ale wth, Ly N ea Ee 29 46op bdahe nee, Moa rinks Ts plslngre Fe ne are

200. Acritosia and Richthofenia ..... ........... os aii ie ey ae Perere: 201. Acritosia ......... Rv, Fa ee ore Te ee, CTA AMO RSs ema REG 202. Acritosia ......... BF a Beige! shea etait Shale ythns deete atte garage Chis Sethi, Rohe ed ete 205... Acritosia. and. Teégultferina: 20200 5 ty. taet ees Pee hk get hes Bas meat 204. Teguliferina TAT Ee PR AS Aa tA inh ae en cn Wy ae ne Ain Pe 205. Teguliferina SSE bt on Mahe te era Stale ott thing uhontarae inet ah ely aes 206. Teguliferina tye = epitome: ta alenglcg Bop gig pale te tS Witarlle oe aubs Gb spate Lule ein 207. Teguliferina iz po ioe a oa eR Sag et AD aye Tet Ree toll EA asa eee gly

208. Paucispinifera, Liosotella, Cooperina, Glyptosteges, and Horridonia .......

209, COOPCHAG se. ag rm Pi, Rig a Ree Ba Aa Wee ge BE. Later ae

BID: COORG gad aay heey oir oe haar Rte eae tte rae Mane ae et ata tet ees : elt: A telestegasttee 2: Med a tFre LS Pry rub ema act ee SM adn ER a terettage 8g Aes = 212. Chonosteges and Cooperina .......... 0... ee cee he eee ies ied Ehhd 213. Limbella |... 000+. RS Pah hee EN POO BE CPR St Ett A re oP Me 214. Limbella .......... Oe eR shtié PA in SR ahted hg Patty a heen eames NE et tlh a ene, ST Aep AA via Cie 2 a, eke se gree a ea re AD Ae NR, OTe TAA E Nk eae dere TO La SIG. Lan Bel ais tise ect BE oe Spo te ee ee le TB, PE te eee he ed tah hee Gas


1188 1190 1190 1200 1201 1202 1212 1212 1212 1212 1213 1213 1217 1220 1241 1246 1258 1260 1266 1267 1270 1270 1270 1271 1281 1286 1286 1291 1295 1295 1297


Page 217, Paucispinifera, Cyclacantharia, Limbella, and Rhamnaria ........ «++. vee eees 1350 218. Edriosteges ..... ... pis its 2 Plnha. aie ant bene ge Ah MES, LAE Oe isa Seeas MERA Pe 1352 B1G, GNIS BOS. Oo ow ct ba ba tales wild, hd edited whee Silhe nah oe eubens ds oe nead TIS OS 1354 DOU. Bdsiostape i. os eke a OE et lace ge eres bed ie lente wae ete Sa eB 1356 P21, Bdriostepes, 5 Shc uke obese AEM Aare Pub Vb Sal Deptt are ach vg NE Sy ube, Eogen dn ne ee ata d 1358 gee. Baviostepes | os Sale toe Fee he yet cn yeta id! Se Ae hay ogee todas be 1360 223. Edriosteges and Echinosleges OS CEs NN fe ses opth pele ee SES OW AWE Ita rd hae 1362 224, Echimosteges: Pocky eSBs eae) ER eae ee pe ee ale Sep ts es eee betters 1364 BOE. ABCRIMOSEE LES 4a Bil Be so alee not ee ch tA tee gfe Whi Eee eh ee ca ee 1366 226) ECRINOStERES song bay ashe 5 Row ces nea be aes pe gldeale <i 9 vote MELE the Had Sarl 8 1368 Dat. EeHimosteges. fot aks! Tw llattig. §alcas eh aderes Pie, Meta eth yo imate aban 1370 S26. Echinosteges. ic. Sec CR Mt WIE) Se a a ak cr eee hay ea ea Se de se 1372 BOG. ECMO eRES LL ot Cena ate ey ale A thug ke Pe ee oe ee eb Spies Oe Pear ae 1374 2300 Echinosteges:. oie bis th emigre oie G hed toes deat s Sak See Mae hate 1376 231. Agelesia, Spyridiophora, and Cactosteges .... cee ence beter bennett gee eens 1378 232.» KENOSLETES =F noone 5 teh hg heen hapeke ge Ses PRRs ome UM are eee rte 1380 233. Hystriculina and Kenosteges ..... 2.6... sc eee cece cen canteen tee eees ve nv = <ESB2 Sat. Kenosteges: 5 a Cok 8 oat LE tu nee oe oe Shad Bataan abey ye ee 1384 285s MENOSE CES eho so Weoley be gle ed a be RW ao breeds ok non ng athe ah ths SB 1386 BOG CKENOSEELES:. Sieh rice Lary Hak | eek Sb ich oo Seeds AUP Seth ars aesing tog Bong 's > ae as 1388 237. Scapharina and Keatorgimella: 5. ou ara ap iE Vapi Sia date Cake alee cere’ Seb gee Pe es mh ouede eS 1390 238: Scapleartna’ vse. ae. pack nse hes shred tel 29 esos eta on ts bE Ceo bw acct see ee lle erat ek ae ge BHO 1392 BED Sea Pavia 8 far Ps lace on sis: cm es He Hy HE ee ABER pi opel go> Te ecie HEE PR aura ele es 1394 240? GHORGHERESS LA a eh tee nee Care erecnty Cog geet yr APM oR Ht me sie & atid sland Sond he Ws 1396 DET GROROSERES oon. - RTT Pe Te Page veel ae Eas, ei dae ean eee eae eed tre CT a ayer’ 1398 PADS CRONOSLETER «Bia Re will Bie. ate ls pee tas, gan eh Rare sg yd Dp ey Mate Pade ae ele Ae EH Bie 1400 285, “ChOnosteges.: 8 ek. sa eee pate Bp de PTE ek EES Scho y eae te = epee ea 1402 244. Chonosteges, Yakoulevia, Echinosteges, Paucispinifera, Oneosarina: and Xenosteges 1404 BERS CROROSEEGES 54 Sa ne ce we ed Nahe Be GH Pa Pn Phe sb arehe Mat angle « gist 4 -. 1406 246. Institella and Peniculauris ....... ...... 0... 125. oh, Me Pd a i SA eee 1408 DATE ristitella* Pee A) So, ee Rian ett airar nicer wats Lines fete te ake ik ore 1 ete Ca AAD ta eat, . 1410 DAS Tr Stitellitry, edhe. heh ge. gh RS Oh oe aE ERE NTS Mine 2 5 bag sesh antace gal ets 1412 PAG PPOs EES | Ne a foe ee Pies aks 5k Sal ad EF ales bv Gn A aed eee See Be Becton 1414 ZOO, GU PIOSEE BES, 5 os a Pare e PHN IES Had Handa re ed ame hee Seg ah pathy rate ey + begs 1416 QB 4, SEVP NOT a3 rw thy Bs AP yee nage be pa - Date eld aim Os ge g Vanes Si cied es ata ieee 1418 252. Spyridiophora ....0 0 2... cee eee Fe, eye ey i nn Le a 1420 253. Spyridiophora and Glyptosteges ........ OE APA i eae eI 2 ee 1422 254. Crasbedona. oo ial \ Scie Pee ree ee ae Re ehh eee yy gu aied ne 1424 255. Agelesia .......... sues Coa te Oho! pose Bat tap cool agit Ng ae lg htetbal Sopra i Mpve BLO Sate TH Sey Et a 1426 256. Ametoria and Rumnventes a, Oe Ce Oe ate atte pene A i Ra 1428 257. Limbella and Ramavectus ....0 0.0... c cc ce ce cece nen e eee eaee 1430 258. Rhamnaria, Ramavectus, and Echinosteges .... Dd Oe eno Ri eB ae F., 1432 259 RRQMAGNE hc. ih ao Odea ara Eben th ais acorns se pS RR ay ese aT he MUS Oe 1434 260. Rhamnaria .... a PAN SP NO ee ON ON, SL PN ce Oa, Ss. en a a a 1436 261. Rhamnaria .... 0 ccc eee Bo sinpt tc ncchee 4 ingly SRR EEE nights Mats 1438 262. NAM atia: Fe boar + Lice en Bee AER do Chetek Beg etd Pega Pe sda peal then eye: 1440 DOS RAMENATIC Wis, ok, ae Eke ee Oa Ms Eee en, Sits a ee Role abt Pay I cere ec Leal 1442 LOSCR Nanaia: © Sore SIR A Ed EES, Atal Bape aah eck ee lds GEASS ahs ime: 1444 DGD MTU GIN ATIC mg Reto the ee ery Oe coe OE UE A sy eile wy 2d Ee Ee oe 1446 266. Rhamnaria ow eee ree mbl ict a Aton eide Sheik me genes 1448 267. Echinosteges and, Ram nari fi. 05 sds oh ee eRe ad gh ph en Bs Bond bab hh Ed Phe dt 1450 PGR DSCHernny SC RCW cee ag he Pim an a BP ts ade fash st og aa cade ce oMiots ya cety be ea Lab t's 1452 269. Tschernyschewia and Strophalosia ...... Mor othe Paces oats ae ere nes eva 1454 270,cScacchinelia, 2-5... cbevt ns st. Sty, Paw h nets tate Ce dade wn OR EAS Se Fae ot ari sas 1456 271. Scacchintellan vac. cabs ody Rite Cs ate ss ess alee eateen Jot ere ty rice 1458 2712, SCRCCHINEUE ec sey so gos Saari ie Sk mAb ele sandy weno els Fay ot ES BS EL ante CS ane 1460 273. Scacchinella. . sc cis ek civic tact erence chester Ae: tchlaloms wk ah Poco aries ae 1462 DI SCACCHIMEa Conse o1 gy faces 0g ts cA eee ale Rela ate oe Bhd ae oe Set 8 dd vara etapa & 1464 275. SCACCTNIANG es hii: srpcs Ge oie AeA ah dhe he dk be BR meee Boe ALN te gith yee bees 1466

276. Scacchinella .... ....... 00. 64.. yt bE TI aah ewe Ae he heen egg Se 1468



278. 279. 280. 281. 282. 283. 284. 285. 286. 287. 288. 289. 290. 291. 292. 293. 294. 295. 296. 297. 298. 299. 300. 301. 302. 303. 304. 305. 306. 307. 308. 309. 310. 311. 312; 313. 314. 315. 316. 317. 318. 319. 320. 321. 322. 323. 324. 325. 326. 327. 328. 329. 330. 331. 332. 333. 334. 335. 336.

Scacchinella ............... Scacchinella ......

Scacchinella Scacchinella


Hercosia .. Hercosia Hercosia PERTEOSUN TY... oe Dies bekes wlae Furey dP pe ahaa sh code Canine Cua Ehaise Egeet Hercosestria and Hercosia —................ ae ice: Hercosia and Hercosestria ................... Hercosestria ... . ...... FE ORCOSESEFTR hi Po ge kg tad Tbe qu ters we gras Pe ora ae Hercosestria ..... Lear EO Ret ty Cee a am ht k ets: Acritosia and Cyclacantharia ............... ht a rae Cyclacantharia. © 06st tie see ies SA fe Sees Cycincanthetia? 02 0. ees nae eee we aed Gyelacintharia ) au oo Brio Byes pi,

Cy CLACAN ERATION, 8 space aw ka AP ney i we 1 ee eget Poel eel Cyclacantharia Pt hd g Ba ce Sele en le, lg ye ne. CRCIACAREN ATI Nc Te Pion cd na eeeta aly des ects abe thath dhdeapu Hercosia and Cyclacantharia .................. eB Eos eae CyCHICONERMEA 0% Rens s AGA IAS DT ROAGE ES Das AD Alwar eats Taphrosestria ....... AE Te SOE a Pind sere ee PE OPRTOSESITIA riot Pa et RG hE Bee ENG a! An oe Taphrosestria and Collumatus .. ....0 1 0 1 ee eee eee Collumatus and Sestropoma ....._...... eh tees hard ee Sestropoma ...... Ps Bhat Lome athe cere Le auto aie unt Acritosia, Cyclacantharia, and Sestropoma ...... 1.1... . cece ee Acritosia, Hercosia, Teguliferina, Collumatus and Cyclacantharia Fimbrinia, Simplicarina, Oncosarina, and Costispinifera ......... Pimbrinia and. Rhytisia. oe a Cees ee A ee feet a alae es Kozlowskia, Costispinifera, and Simplicarina ......... .......

SICOZIOWSRUR: 5 pS Re erizeew avy 0 5c tic Seg ae Beem A eG deg clea

Kozlowskids ok re ee bt aces cae cee ate oat Oe ete Mystricilina ©. esl, gi ha bony Pare Lanier, Sh ait. Bothrionia and Thamnosia 5 te, Ae PS Bothrionta ao. hoe Se, eA Lene eae... Sire, Oncosarina and Bothrionia ..........00.000500 cece ee eee GOstispintferae nr ,. cek bth ots aed eee ghee PPagaed Costispinifera 66. eee Se ut eee ese Costispinifera me peach eh PSpes see, Wendy ioe mpaiiee of Eliliotietla: *& ac. (ytd ood Ae Say eerste Co ee, a, Pr FLT OLL CL bea metre ps, dys tp eed yee te elite DY te foe ee Oe Mee pea ae OVICOSGTING: Mes 8 cs Ke ete re pole ed ek Ea Be Hystriculina and Elliottella ...................

Echinauris ...... Fe ee Ee ee aE PEN. BO ee eee Echinauris. Foe A Sirs bean tee Mae Ree Seg | pa ee: ECRINGUTIS: & PBs. 4 casts ods shige Buin Bethe BO Qe tk A. f. .5.0 5 Se Echinauris .. ee ee eS eee tee ee ces Echinauris .. .. a Se 2 per ae

Echinauris ...... bled are 2 Wey Gee

Echinauris .. Echinauris .. Echinauris ....... 0 ...... Mi, ARR Awe Fo, pee Echinauris ..... al ate. ¥en nase BERiNGUTiIS=. ee A Ta ae op Behe age Ale eS

‘Scacchinella ..... spe eet CAIUS las 8 +e ty i

re oe ee ee)



Page BaF ECHIMQUYIS. NS hg Ate Pp eins eileen gdalrttes gab tahun malic : Oy eet ccd £ ata’ o 1590 Bod, DCN GUYER ON, Fy Peso yi Be, cet it Mh She PER a te Ad ba Uitte, Penne eR eares 1592 339. Echinauris .... A SER, eos © ae thes PaO Ee LER. Se On So 1594 340. Echinauris ...... Ft Ae CP CO Pe ee Weer Eke Nae ee te nt Rede oem .... 1596 S41, PCRS Ae 8 sly et eB Oe EB Ae EGS le SE Qe hy orn dele, JoB RG oopaetin eh 1598 342, Kutorgivtella® oh. 6 oc udeekanbe nde te dyes te pe PoP y Mie fecstich oa yas Oe ee greedy Bee 1600 SOG CR ULGTRENELA 224-5 hulad Be ee ins Eh aca oth 4 PB Ret ech inet gh pl ae een es 1602 Bad, Kestorginella: No cogs totes th ele he ee dy ee pcteags t netie hid feiding 2 anaes 1604 B45 ERAMNOSIA, 2: Nb decd ln nee oe a Gh nage OG Sead ale ~ 1 Seat eth es oO Le eae 1606 BAG. ET RAM MOSta nen gga. Gley oN ecacane De PRL Whee nh eottagls ulead OD las Rig edhe TE eat oa 1608 S47 eT hamnosia” 2.) 08 yore ae OS ST My ea omendepimaieeuats abyss oe by wie 1610 348. Thamnosia .. 0 ce ees Bote in: 2 oe a, Aaa eee See 1612 349. Thamnosia .......0 vee eee eee eet etch de nshdleg xt somiochinhe e ealas Beets 1614 S50,; Kutorginela t.-£5 song den es pea Shs - windeep Aa thal shit SI ihey bea tee bie: Ore aes SF 1616 351. Thamnosia-and .Rhammnaria: 2.505 ee She bag ona tie ee tg hae gy eee 1618 352. Bathymyonia and Waagenoconcha ..... obit codhupel phere te sty loge Et emote BPO Beto, ciao Pay Tea ay 1620 353. Hercosia, Juresania, and Bathymyonia ............00 vce eee eens 1622 $54. -Wikeeencconeha 9a. .cc, UE na PLP APA AG ete re Bye thereon hes Was aeue lh emcee 1624 355. Waagenoconcha ..... ©... eevee eee eee i alt oy og, 6 ee at tee Bary ee hte : 1626 356. Waagenoconcha ........... abt dal de or anigteceiaAd pot get ane athe MOMS Wail ich sett eed 1628: SETS Witla tenOConER@ «farotsdavbel Cored PISb ee dc UHghe eet beh ale fe HORSE CEA frie 1630: 358. Kochiproductus .. .... EMP th eel OAS MR asd Vie ete tee ORE, | ee arlene 1632 SHG. SIKGCHIDVOGUCTUS 214, 2s. reer pa: Sten H 29,0 Re HE er aE & Fan Cotlataees Meh eng ete ML 1634 BOO. ROCHEOVOAUCTUS: aa eye Cag tts tbat ea ee a ROE PRT ee eal ale eal a Pee eye EE eG 1636 G1} FOCKIPrOductus yo. ac. ek ID Ae eee Fg hg Og Ee hal Pleas ie ait SERGI RU theme 1638 362. Rhamnaria and Kochiproductus .............000 ccc ccc eee cee tees .... ©1640 363. Dasysaria ...... .... Sa eer aah gd wee a a tae he neste toe en. harbor Ute Oe cel ea ta ote die niki 1642 REFS STE. Cl aioe SMe Huns hae OS See NEM EER aE ed ape PA badge Sen ete ap danos ee yak phe te 4 1644 965. Dasysarid= dnd NudGUris: oie been ace er ak Des Appa th Bd ent ee help Ee 1646 366. Xestosia and Nudauris ........ 0.0. c ccc cc ene e eee e ee bene ae 1648 367. Echinauris, Xestosia, and Nudauris ..........066 cect ee ees BR sed teat all 1650 BOS INUCAUTES (oo He REA oe, Gad Whe oe oe it mal hut me. Bible ca | tet tye lala, enue, SMe ee tans 1652 BOSe NUGAUTIS. Bone hee Per ae ne A, Se gt Nala, eee fe td de eating yo dots MEPL ESE cour aint ¢ 1654 STOMNUGAUTIS. SS occ: ea esartes oe te Pe eed adage eae PUA hak ats loch ws die atl AMA ESE Ser 8 OE ale 1656 S71. Nudauris: and “XestOsta. (14) sary gabe ed iho AAG OSG eel eee dod ge tee Bs oe bah 1658 BPP INUDAUTIS a ecec B Shauretnc de BONE apechcdtnp re Meee ence ta ak Pate MONE Pkt tech cll tee NOR eoartcl, Sob ceateag tee sues 1660 BT et RRO ice aesen dF 0 45 tay ed Eee ks UE DEC MITE AUN OE Vases LL erature gah hee eset 1662 STA RUG AG 8d ce | ep yates tan Ae hy Shei Bae, Spates ty aya og PT ERAN fo ah bud ciel bat aA g Eira 1664 375. Rapala Mig ela k gett a aot v ADR ANGE dip hs es A eR a cB ens * 1666 BG; PUA th NN, ene af ols 4d poe 1% on poy Eke eyed found ota eof SPAY oleae neck fever eates tes 1668' SUR Regatta BS ees Pela ei a tale bead ents AEE ohare acer ace Re Ne ean ag errs 1670 a(S: Rugatia and Reticutatia: 4 ccs eee ek eased ide en tse kien ds Mas 4 het biayetG 1672 SI, BPINGTEMG, P07. i2y. satya gS era key Re SEY ROAR EN CE IVI Mig SORES Ms TE 1674 SOD PTOI ELLE «am BAe evita hoe srt eee een ce end arb, Mets (I> fara nt hci, dave ose 1676 SOI SHAVE Ace o orf ok TE Sr stete neat techy, setae hare od dts Bl a nae phe a trap ee yee ATEN 1678 SOLS KOSLOSEG soi A Se Mar 8 ag wie hs ee ache De Meh, eke g hese recto Medes ole te hind Day sade ere 1680 383. Xestosia and Reticulatia ....... 0... ec ccc cect e en eeteneneean 4 2. = 1682 Set eSpeiniirons: Ost ot Pee Bees Fer Oe hee be Sea es ed Pe APRA a> derek ... 1684 SEB. BP fons os Ae co eB es ts AP hee TOP eee eee s 4 Sak AN sxe Seetese hog Ase 1686 © 386. Reticulatia, Antiquatonia, Calliprotonia, and Echinaria .............. ws... 1688 BST EPL ONNE fe Sie eh ly Beh Fe Ae TATE fae DTN Ae pitch attats ae no etenn hace peed Sey - 1690 SRE ASP Herons ct cco ce ee bate & eee de Le ROL Rd Sd ncther Mies Be Nah then States . 1692 SOD SPAT ONS, nan M sent oe Dah hs AA ed py OT RHE Rs eee s ROA. win chs, Meslatelradt ot -. 1694 390. Spinifrons and Antiquatonia ....0.... 00... cece eee eee eee aes athe 1696 BOVE Are taGUAT ONE secede pinse Be gn ste & Apri ada adeeb edces fa Ratner ental, sess 1698" HOD, eticmlatia: 650i. Fy4 3 il. oes gran ey bea yao deste hao reine etn ater EAPO NE an Sg ih he 1700: 593: Retieiulatiae suns Bias 505 Sa Nes Gn pon 4 SaePades te Bag se Po ede bu bate ah Se, 1702 $94 Penioulaurise | Cn «sea tee nes oe his Bataleon hE Bee oees)©61904 395: Pentculauris. cue. vp hale ARAN Lea WROTE SL ea ei lnal caw niet Oe be ook el glee eans geal .. 1706°

BOG “PEnicilauris Bye Ce Ge A: UN belek hR aye oe ee Be Papeete te in oes we 1708.


397. 398. 399. 400. 401, 402. 403. 404, 405. 406. 407. 408. 409. 410. 411. 412. 413. 414. 415, 416. 417. 418. 419. 420. 42), 4292, 423, 424. 425. 426. 427. 428. 429, 430. 431. 432. 433. 434. 435. 436. 437. 438. 439. 440. 441. 442. 443. 444, 445. 446. 447. 448, 449. 450. 451. 452. 453. 454. 455. 456.

Page Pentoutauris” oo A Apel ete D1 | Maatat tte sew oat 1710 PENCUIAUNS A ie an os doh trace ttt ©. asthe. td, Bin MeL ee 1712 POTUCHIAGTIND Gia ls hae eh tee eae eur et el eke a tha: Tacos tee ele 1714 Pentoulauriss 68 Aah ee Bisset eae es yl er ie a Ed 1716 Pevisculatts on I i i Ne re ee ate ot oR ORO, Bone ce LEN, outa 1718 POUCULAUHES or) sigs ph, ncectae eee eeA ae nol) | Use eae 1720 Hrerincte lari Bn! ey ee hee nate! Boke p oe Me A een ato) the ei tats 1722 Perectelae rie TS 8 Ps edocs, lean is gh meteors A osoak vith ae ea a al hy gee bee Betas 1724 Penictlauny =... Te See Sea 2d de WAP ALE a Ee NR) bah Lo eee 1726 PORTE TATE =. Lt hide «dete BY Gon Tras ore Ee tk lets tngs ate OAH IS babe de h danGte 1728 POCUMAUTIE a eos Ast Mute ee ein ee htc ete Mutat ae a aypedies Aap Fee ee: 1730 MUOTRON GILG cose Sosa s noite see Px Tes Ee RA ET Bas ea de ee es eae he Saad Bos Fea 1732 BS 21 le ee SD Re ee Lek tees ke fetid Nh, Pee ine | FORD WN Ki bale 1734 Nudauris, Echinauris and Liosotella ..... 0.000.006 cc ccnunscunevvccvcetcennceccs 1736 EAOSOLOUIE Boe con nets cote yah cee en Eee el gana eee LAR DEP Sa 1738 Liosotella and Grandaurispina . 20.0.0... 6.6 cence cece en een ee tep ene enenenee 1740 Liosotella ........ sade SPE ie tees ele cP ties wo ALE ee ae as OL eine cae ae at cata cased Mame ad Pcl ats oe 1742 TAOSOTO GS on A lsh tenes New AOR Net, et, ee, eet a eee oe lh el Ny 1744 Lvosotetla- 2 ht A ee ER EE ok Dg eat setae tees Mors Ahr eeteeeee a dete toe 1746 Eiosoteilat Ak tx, pee we kek ee ee Se ee Sa SE See ae alae cur >. te 1748 Paucispinifera ........ .... EE ger EL NTN ree aCe Ot EON I nsf Wea RR 3 1750 TUGISPATETOTE. Ens becca tek Boh de De Cah. shy cs ie, Sat o. Mege th TE Rage SOM aah ary 1752 Paucspinifera and Oncosaritit 2.005 odds ee - alight oP ive Cus llun os ee ctbeRaty 1754 POUCH PIPAZONE 6 in worn snd eb ong ope gs wea SER as aoe tahoe WELD LALA SEE TR A 1756 Padeispinifera: 60s 60 diiages Fas dvdr Ah ae $4 Aoiiattin Fetchaa hs 1758 POUCA OTE. 1 Awcy Jobat. Bim ale Wits c Ae Pome, | pele k Att Ere a te, .. 1760 POUCIDTD ETA: PI hod i096 cece dc Bs aie On ee | ER eee Ute yA Gee te Swat 1762 POUCPIRIETAS see Daa aoth Se eens see ee Pn Pree cir . 1764 PAUCSPIN ONE. oop aes os Wien tee igs mace FE a katy Spin gen 4 elec ae 1766 Paucispinifera ...... 5 Gabel Spgs oh te ee dest bee a Ne eases 1768 POWWNOPPROTIOS 6.55325. 5 ook ho BSc abe Ds ee ag whew ; 1770 Gancrinelia® 6 ac thier ed che Ka ly eats oe ES 5 SSeea., 2, See ee Ee 1772 Cancrinellas oi. phi) pd th. Nae NO ee oy 4 a eed Me 1774 Cancrinella and Grandaurispina ......0 1... cece cece tenet eee et ene teens 1776 EAR ODTGUUCIUE. We bole ae Ren Oke A ee ETP pee tbn Ro Peres 1778 Linoproductus nals eeceacst ese ea pee tye a Deh et bers & a4 1780 Compressoproductus and Linoproductus ...... 0 ....... 0.000 cece cece eee 1782 Yakovleviav and Ltrvarianc. oh Ack es he tre h pim sch Eite. DO 2 EE atgotese's 1784 SPOR A RUTISHINA OA FP 5 Fcc ity) Aye eee he PARR RS LAH bocgotce ee ged RAR ICE ee Salar heb ois3 2s Bey 8 1786 Grandaurispina cee nee eee nee eee pg vee od sheet Tis OeA Ba pActeias yo 1788 RSPR UTES RIT Bo ik a EU wet eR OS ey he ct este aleve oh Su aly atest 1790 GHORGGUIIRENGE 6 Fra Be ot 2 Bs stu ety Bag gh Regn oo Obes hber oyna titer as eet anda 1792 SSHOMAQUPESDIRG oof igo cece g te peep ein Ph | fae aay eed peg Meee pe EE 1794 : Grandaurispings, 246.2004 5 8574.5 Shen epee Beth ip TES OPS ORG Mad be Ma Oa ep ae ae 1796 Grandaurispina On ie LA RO ce - in PRS Fy eee mn eee oh raya EF 0 1798 Grandaurispinad ee ce eee TL". ee hy: Re eee eM care et; 1800 GranGaurisPingy 25. ls ae 5 ih sy oa png ees eee Dean kd ag ton ahs Micha ara de 4 1802 Holotricharinaa «io. Me a, meee are wie elas ahha uae Peateoe ents 1804 Echinaria, Holotricharina, Hystriculina and Calliprotonia ...... ........+-.055 1806 Acanthocrania, Paucispinifera, Elliottella, Liosotella, Megousia, and Holotricharina 1808 Merousia ‘and. Kovigwshig 806. ole Ads tad LAOS E LO Reais tek S ee 1810 Megousia ...... Ree Sif eked tedewen plein a he Macatee Marten eB cone ots ctare ok wg ata ny A 1812 DAE EOURG POM feb MO Re Re heniietgt: Nels SRS Ae Pe Rekecty Rabat ok Park aaa GP Maceo! Ma clas dy 1814 IE ROSIE Ue Seg Sica co mufenen es 1 best evig ere aft edeug tan VR dco argcanaceth Sec maak Page abe rao 1816 DIC EOUS ia, de | eu ries 1 Mee me ie || OM) RAL PE Betas . 1818 Megousia and Yakovlevia ......... .. RET epi es bee trees Ok a a 1820 Megousia, Grandaurispina, Linoproductus, Antiquatonia, and Kozlowskia ........ 1822 Compressoproductus .....-. 0... eeu eae oe eer alerts TEP ape va Serer ei 1824 Striatifera and Compressoproductus ...... 4; a ial se tee tea bees bee agg 1826 COMPTESSODTODUER ES oe Be) ope nS dpe tot bg eta ~~ boerapethn «las dpe Je Deb eee es 1828



Page 457. “GOMTDYESSOPNAGUCEUS 1D cA acoe beds tink CART AT AS anes Std Eee ctbey dolce poe .-. 1830 BBB: GOmPprecsohrOdwetie xt. us) sce yes pee ee baa te oe reel ee Lek Pee er ek eo a 1832 459. Compress productus. al iceyh, cuts Seg pedo oat Ute apgk ak = dee dda sed tees ep cleans 1834 460. Compressoproductus ..... ic cece ccc eee PEMA S ores SS ee Rie LORI 8 a’ 1836 AGI, Compressopradiictus: (2) oe cae ee ten eye tein ate ons Dhak de PADMA NE tole es 1838 462. Compressoproductus and Striatifera ©. 6... 61 we eee eee eee 1840 463. Institella, Megousia, Compressoproductus, and Striatifera ............0 eaves 1842 464. Compressoproductus and Polymorpharia .. ©. 0 1... cece ee eee eee eee : 1844 SO CSET ETA we Lt le poe ep ards ig od PG AL Drader ads lore ee scl Galt atlfig yp ek OF peat de age 1846 S66.“StDhOnostar so e/cice cee At ace tee Na A Ee Pte aA g Pa est alte alge RAIN OR OAL Beale «Fog ah .. 1848 467. Siphonosia, Megousia, ANG ACHAEA Mi os, cnn So een eared pe ME Nas Ne . 1850 468. Horridonia, Echinauris, Institella, Glyptosteges, Siphonosid, and Nudauris .. ... 1852 SGOS VGROuleUia ne ne oe Se hag peso Pantin aa Ms See eee We re Poh tart Marist 4 1854 BIO Vane lean as 655 co ho fe Greg Tae ov As wince o/die atey se els ng HED els ph ae A VO e Ad cb ares 1856 AT. FOROUMEDIY aio SBS EVE nee thle MNT OE A oe CP RNs red Said Ode ete’ arts 1858 BID) VORQUIGU Aaa oO onde nila erent ed seta hd bibs bet lap t Welty Rate See Hite Bice CBee sib 1860 473. Yakoulevia 9.00.0 ccc vac ew ete v eee eee pees CAA, rae ATS AE Rae em! 1862 414s VaRouleuta. Rhee ohn Sedagra waded Gouna sileas scene fs ead Nake 1864 475. Spyridiophora, Paucispinifera, Limbella, Striatifera, ‘and Orthotichia ...... .. 1866 476. Echinauris, Sestropoma, Paucispinifera, and Rugoclostus ........... ..++.- .. + 1868 ATFs GROTEURERES. dis Mie pone ice ts wed Aes by cle wenans of ie ae WAT AN Ge Bs eed lena . 1870 478. Chonetinetes, Chonetinella, and Sulcataria .........0 0.0 es Eee 1872 479. Chonetinella and. Leurositia’ (o.oo einen 4 seek hn ts HA a Loe renee See oe elOEe 480. Chonetinella and Sulcataria cee eee 1876 PERE PV OTOS: LD OPIS) Be aa eng rainy Bee ach Gee Sed mph ead a Rohwer fia phn yn Catala oe 1878 482, Dyoros (Dy0T0S). 2.0.66 een ewe thoes Pie oa heptane wtatihpens Drag pe = Reet ps 1880 483. Dyoros (DyOTOS) ....... cece c cece eee oe L2G BS Qe ee OR ea ae 1882 4840 Doras YDVOrOS)} Oy ss 4 cg ergy igs A tig Mee Png th co Mee halen pets tate ane oie Al el Stabs oy . 1884 455. OOS (DyOrOsy. ot ps Pe REE nly Ads POGUE SEED BURA Te Rtitong Lee At gla es £43 1886 486. Dyoros (Dyoros) PI eins Se te SAE ep oe EO EAN ee Ae OR LIT, A De Ree ee 1888 487. Dyoros (Dyoros) and Dyoros (Lissosia) ..........0 ce cece ccc ee pee eee 1890 488: Dyoros (Liss0sta).. occult Matis a gts nice decane ah Sa ic rec trace a eed iy OUBAE det Tankers, vag. 1892 489. Dyoros (Lissosia) and Dyoros (Tetragonetes) ... © .0.. 0c ec eee eee eee sees oes 1894 496. Dynros (Letragoneresys ix Sop oiah baa te Gd tye hen ae he ep aeray ein Rng sa sitptes ste 1896 491. Dyoros (Tetragonetes), Lissochonetes, and Quadrochonetes .......... elias 1898 492. Dyoros (Tetragonetes) and Undulella .. 0 6... 1... cece cee 1900 SOS SMECT ODEON cy wedi nltg © oc ba fk GS ep mg ly died fy Meats Asha 4 olaels hue a Eg nls een 1902 BO4! ERUT OSM G5. cl ocgrce tn cc kcbocca yma giaitefe pA, Lf ig 8 OPE aOR KS pale Mes CA OES eta 4 OE 1904 495, Leurosina and Dyoros (Tetragonetes) 0. ccc oes cee ee awed eee eds hes 1906 496. Mesolobus?, Rugaria, and Neochonetes .... 0... cece eee tee ee -. 1908 497. Neochonetes, Dyoros (Tetragonetes), and Lamellosia ................ 0.0405. .. 1910 498 Rugaria, Leurosina, Dyoros (Tetragonetes), Dyoros (Dyoros) and “Chonetes” ....... 1912 499. Dyoros (Tetragonetes) and Rugatia . 0.01000 oo ec ee heen beweebabee tes 1914 500. Chonetinella, Dyoros (Dyoros), and Leurosina .... 00 0. cence eee 1916 501. Dyoros (Dyoros), Chonetinella, and Micraphelia ............0..00.0.000 0000 ccc eee 1918

502. Quadrochonetes, Undulella, Dyoros (Lissosia), Dyoros (Dyoros), and Chonetinetes .. 1920

Permian Brachiopods of West Texas, III

G. Arthur Cooper and Richard E. Grant

Order STROPHOMENIDA Opik, 1934 Suborder PRODUCTIDINA Waagen, 1883

Superfamily STROPHALOSIACEA Schuchert, 1913

Family STROPHALOSIIDAE Schuchert, 1913

Primitive Strophalosiacea with hinge teeth in the pedicle valve and sockets in the brachial valve. Pedicle valve having an interarea; delthyrium closed by a flat or convex pseudodeltidium. Noto- thyrium, if present, covered by a chilidium. Pedicle valve anchored by rhizoid spines and umbonal ce- mentation. Cardinal process initially bilobed but becoming trilobed to bulbous with age.

Subfamily STROPHALOSIINAE Schuchert, 1913

Strophalosiidae having a smooth dorsal valve, without spines.

Genera in West Texas: Strophalosia W. King (1844) is extremely rare in West Texas, only four specimens having been found; Heteralosia is a genus common in West Texas in the Permian and Pennsylvanian.

Muir-Wood and Cooper (1960) erroneously de- scribed the subfamily Strophalosiinae as having a spiny dorsal valve. They then created the Hetera-

G. Arthur Cooper and Richard E. Grant, Department of Paleobiology, National Museum of Natural History, Smith- sonian Institution, Washington, D.C, 20560.


losiinae for genera of strophalosiids having a dorsal valve without spines. Brunton (1966) studied the type specimens of Strophalosia gerardi W. King, type-species of the genus, and was unable to find any trace of spines on its dorsal valve. He therefore recommended the elimination of the Heteralosiinae. Heteralosia proves to be different generically from Strophalosia but belongs in the same family.

Genus Strophalosia W. King, 1844

Strophalosia inexpectans, new species

PLATE 269: FIGURES 13-30

Medium size, but large for genus, wider than long, subquadrate in outline; concavo-convex; sides mod- erately rounded, greatest width at midvalve; hinge narrower than midwidth, straight, with short in- terarea. Anterior margin rounded to somewhat truncated medially. Rhizoid spines short and deli- cate, concentrated on ears, umbonal, and postero- lateral slopes; ornament spines numerous, delicate, recumbent, covering entire surface.

Pedicle valve fairly evenly and moderately con- vex in lateral profile; anterior profile broadly con- vex, moderately steep lateral slopes. Umbonal region swollen moderately but extended slightly posterior to posterior margin. Median region greatly swollen, long anterior slope moderately swollen but not steep. Interarea orthocline. Del- thyrium small and narrow and covered by a slightly convex pseudodeltidium. Ears poorly developed, obtuse. Cicatrix of attachment small.


Brachial valve not well known, moderately concave; deepest posteromedially; posterolateral regions flattened; interarea short, hypercline. Or- nament spines thin and delicate.

Pedicle valve interior with small, delicate teeth very closely spaced. Adductor scars large and rounded; diductor scars flabellate and surrounding adductors.

Brachial valve interior with long narrow cardinal process extending nearly perpendicular to posterior margin. Sockets deep. Adductor scars elongate; me- dian ridge low, rounded, extending to base of car- dinal process and to about midvalve in anterior direction.



ial sur-

valve _ face mid- _ hinge length length length width width height

AMNH 512 151229a 26.9 23.0 33.0 30.0 20.0 10.6 151229b 23.6 20.5 28.0 30.4 22.0 9.5 (holotype) 151229c 23.4 19.4 27.0 27.3 20.8 8.0

STRATIGRAPHIC OCCURRENCE.—Cherry Canyon Formation (Getaway Member).

Locatity.—AMNH 512 (= USNM 728).

Diacnosis.—Large Strophalosia with numerous fine recumbent hairlike spines on the brachial valve and short rhizoid spines on the ears.

Types.—Holotype: USNM_ 151229b. Figured paratypes: USNM 151229a, c, d. Measured para- types: USNM 151229a, c.

Comparison.—No_ other strophalosiid of this type is now known from North American Permian rocks. The genus Strophalosia, sensu stricto, ap- pears hitherto to have been restricted to the Perm- ian of Asia and Australia. Some species have been described from the latter place and the type of the genus is from the Himalayas but it is poorly known. Species from Australia usually assigned to Stropha- losia have a spinose brachial valve, a feature shared with S. inexpectans.

Strophalosia inexpectans cannot be compared favorably with any of the Australian species which are much larger and more robust, usually with stronger spines and with details of the interior different from those of the Guadalupe Mountains